It really depends on your area of interest.
For example, you might want to translate documentation, web stuff on the site for your distro, the installer for the distro... In such cases, you want to get in contact with the people of your distro, there might be instructions in their web site about how to start.
If you want to translate a given program, then get in touch with its author upstream. That way the whole community can benefit from your work, and not a single distribution.
There are bigger projects. For example, the kde people do have their own translation team, so if you use kde you might want to join the translation team for your language, or create a new one if none exist. I guess it's the same for gnome.
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