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Are you sure there is no slash "/" after mail.mydomain.com? If that's the case you should get a "Server not found" or something like that because of the wrong domain name (mydomain.comsrc).
Getting a 404 error means that the domain part is correct and you're trying to access a page that does not exist. In this case check the web server error_log to see what this page is.
Firefox can't find the server at mail.mydomain.comsrc
In this case it's not a squirrelmail misconfiguration.
It looks like you're doing some sort of URL redirection and you have forgotten the trailing slash.
If you're running apache check in httpd.conf and/or .htaccess for the respective directives
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